Wildfire has fundamentally shaped the western landscapes we seek to conserve. It is a source of renewal and central to the functioning of many ecosystems; as well as a destructive force that threatens communities and conservation values across public and private lands. Wildfire epitomizes myriad connectivities that we cannot escape. Yet as the frequency of large, severe wildfires has greatly increased over the past 20 years, it has called to attention the fragmented, sometimes conflicting approaches to natural resource conservation across different jurisdictions and organizations. Reducing threats and enhancing conservation benefits from wildfire will require synergistic collaboration and coordination to span these disconnects. Our discussants will provide a range of perspectives from applied social science and policy action around wildfire to suggest bold new ideas about how people in the western U.S. may live with fire in the 21st century, and how conservation policy could spur more effective collective action to address wildfire risk across public and private lands.